Paradox of marine protected areas: suppression of fishing may cause species loss

This is a simple model of a prey–predator system in two areas, one of which receives fishing activity (fishing ground) and another that does not (MPA: marine protected area). Takashina et al. 2012 gives the following description of the model. "A number of fish and invertebrate stocks have been depleted by overexploitation in recent years. To address this, marine protected areas (MPAs) are often established to protect biodiversity and recover stocks. We analyzed the potential impact of establishing MPAs on marine ecosystems using mathematical models.